Motivated by last year’s fern forays and spurred on by my new membership of the British Pteridological Society, I have decided to try something different this year: a personal challenge to see as many of Britain’s fern species as possible in a single year. The idea being that, by seeking out our ferns, I’ll learn more about them, their habitats, and the subtleties involved in identifying them. This blog will be part #10.
Teesdale – 31 July
Friday (my birthday, no less!) saw Matt and me heading to Teesdale with several members of the Natural History Society of Northumbria, our aim being to take a closer look at the plants associated with the area’s crags and screes, ferns foremost among them. We spent several enjoyable hours rummaging around the rocky slopes, encountering many of the species that characterise these diverse, if somewhat inhospitable, habitats.
Among the commoner ferns, Lemon-scented Fern Oreopteris limbosperma proved far more abundant than on my visit earlier in the year. Parsley Fern Cryptogramma crispa was also present in good numbers, now showing a pleasing mix of fertile and infertile fronds. Ever a favourite of mine, Beech Fern Phegopteris connectilis was encountered in a cool, humid spot beside a small waterfall, where it looked very much at home.



Continuing on and turning our attention to the boulder screes, the first ‘new’ fern of the outing soon presented itself. New, that is, for this year’s challenge rather than for me. In fact, it’s a species I’m even attempting to grow at home thanks to the Pteridological Society’s excellent spore exchange.
The fern in question was Mountain Male-fern Dryopteris oreades, a geographically restricted but by no means rare species in these upland habitats, where it thrives on inaccessible rocky ledges and among unstable scree. It is a lovely little fern, more delicate in appearance than the familiar Male-fern Dryopteris filix-mas, with pale, whitish scales, slightly twisted and uneven pinnules and, perhaps most usefully, sori that occupy only around half of each pinnule rather than covering it almost entirely.
We found a few plants at first, then many more, and before long gave up counting altogether.



Ambling along, making a bee-line for the site of a rather special little fern, a few new species presented themselves. We noticed Green Spleenwort Asplenium viride and Brittle Bladder-fern Cystopteris fragilis on areas of limestone; Hard Shield-fern Polystichum aculeatum and several examples of Narrow Male-fern Dryopteris cambrensis. These were, however, promptly forgotten when we finally laid eyes on our target.


I seem to have a rather frustrating relationship with Northern Buckler-fern Dryopteris expansa. I’ve set out in search of it on several occasions, only to return with what I thought were convincing examples before being told they were, in fact, merely small Broad Buckler-ferns Dryopteris dilatata. The two species are undeniably similar and often grow together, though Northern Buckler-fern is said to have flatter pinnules, paler scales and basal basiscopic pinnules that are noticeably longer than the corresponding pinnules immediately above them. Tucked beneath a large boulder, the plant below appeared to show all of those characters, at least to my eye.
As you can see, it is a rather delicate-looking fern and certainly much flatter than the typical D. dilatata. The fronds were also a noticeably paler yellow-green, while the stipe scales lacked the conspicuous dark central stripe that usually characterises Broad Buckler-fern.
For now, I’m happy to call it Dryopteris expansa but, as ever with the trickier ferns, I’m more than happy to be corrected by those with greater experience!


The Cheviots – 2 August
A few days later, on 2 August, it was time for our now annual excursion into the remote and beautiful upland corries of the Cheviots. After six hours of steep, and at times rather tiring, exploration, we had amassed an impressive list of botanical highlights, ferns very much among them.
Mountain Male-fern Dryopteris oreades was once again well represented, growing in its favoured haunts amongst rocky ledges and boulder screes. Having become better acquainted with it in Teesdale just a few days earlier, it was pleasing to encounter the species again in one of its classic upland habitats.

Whilst Parsley Fern Cryptogramma crispa was looking particularly lush here—perhaps a reflection of the greater rainfall—it was the clubmosses that really captured my attention. Fir Clubmoss Huperzia selago, one of the fern allies, was encountered in several places and looked especially handsome, rather like a tiny pine or spruce tree clinging to an otherwise inaccessible rocky ledge.


Much more exciting, however, was a fine colony of Alpine Clubmoss Diphasiastrum alpinum, growing in one of the most inaccessible corners of the corrie. Perched on the edge of a large boulder where competition from the surrounding vegetation was minimal, it occupied exactly the sort of exposed niche in which this species thrives.
This is a plant I have only encountered once before and, on that occasion, the population had been badly scorched by drought and was rather underwhelming. Seeing it here in such good condition was therefore a real treat, and a fitting way to round off a thoroughly enjoyable day in the hills.

I haven’t had as much time to devote to this challenge as I’d anticipated, but that should change as things settle down after a busy summer. My first trip to Ben Lawers is pencilled in for a fortnight, and Matt and I are also planning at least one jaunt to Ennerdale in the Lake District to see what ferny treasures we can turn up there.
Whatever we find over the coming months, this challenge has already introduced me to far more of Britain’s remarkable fern flora than I might otherwise have encountered. I suspect it’s safe to say I’m well and truly hooked.
Running Total
Seeing as I rarely carry a notebook, we’ll log our running total for the year here…
| #1 Wall-rue | Asplenium ruta-muraria | Tyneside |
| #2 Maidenhair Spleenwort | Asplenium trichomanes subsp. quadrivalens | Tyneside |
| #3 Hart’s-tongue | Asplenium scolopendrium | Tyneside |
| #4 Black Spleenwort | Asplenium adiantum-nigrum | Tyneside |
| #5 Broad Buckler-fern | Dryopteris dilatata | Tyneside |
| #6 Male-fern | Dryopteris filix-mas | Tyneside |
| #7 Soft Shield-fern | Polystichum setiferum | Tyneside |
| #8 Japanese Lace Fern | Polystichum polyblepharum | Tyneside |
| #9 Hard Shield-fern | Polystichum aculeatum | Tyneside |
| #10 Hard Fern | Blechnum spicant | Tyneside |
| #11 Western Scaly Male-fern | Dryopteris affinis subsp. affinis | Tyneside |
| #12 Borrer’s Scaly Male-fern | Dryopteris borreri | Tyneside |
| #13 Lady Clermont’s Spleenwort | Asplenium x clermontiae | Northumberland |
| #14 Rustyback | Asplenium ceterach | Northumberland |
| #15 N/A | Dryopteris borreri morph. robusta | County Durham |
| #16 N/A | Dryopteris borreri forma foliosum | County Durham |
| #17 Greater Scaly Male-fern | Dryopteris affinis subsp. paleaceolobata | Tyneside |
| #18 Tunbridge Filmy-fern | Hymenophyllum tunbrigense | Northumberland |
| #19 Lady-fern | Athyrium filix-femina | Tyneside |
| #20 Adder’s-tongue | Ophioglossum vulgatum | Tyneside |
| #21 Delicate Maidenhair Spleenwort | Asplenium trichomanes subsp. trichomanes | North Yorkshire |
| #22 Narrow Male-fern | Dryopteris cambrensis | North Yorkshire |
| #23 Green Spleenwort | Asplenium viride | North Yorkshire |
| #24 Parsley Fern | Cryptogramma crispa | North Yorkshire |
| #25 Moonwort | Botrychium lunaria | County Durham |
| #26 Holly Fern | Polystichum lonchitis | N/A |
| #27 Oblong Woodsia | Woodsia ilvensis | N/A |
| #28 Western Sword-fern | Polystichum munitum | Northumberland |
| #29 Oak Fern | Gymnocarpium dryopteris | Northumberland |
| #30 Beech Fern | Phegopteris connectilis | Northumberland |
| #31 Polypody | Polypodium vulgare | Northumberland |
| #32 Royal Fern | Osmunda regalis | Yorkshire |
| #33 Lobed Maidenhair Spleenwort | Asplenium trichomanes subsp. pachyrachis | Yorkshire |
| #34 Rigid Buckler-fern | Dryopteris submontana | Cumbria |
| #35 Limestone Fern | Gymnocarpium robertianum | Cumbria |
| #36 Sea Spleenwort | Asplenium marinum | Northumberland |
| #37 Small Adder’s-tongue | Ophioglossum azoricum | Northumberland |
| #38 Narrow Buckler-fern | Dryopteris carthusiana | Northumberland |
| #39 Brittle Bladder-fern | Cystopteris fragilis | Northumberland |
| #40 Lemon-scented Fern | Oreopteris limbosperma | Northumberland |
| #41 Mountain Male-fern | Dryopteris oreades | County Durham |
| #42 Northern Buckler-fern | Dryopteris expansa | County Durham |





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