For our next plant profile, we’ll take a closer look at one of my favourite ferns – the delicate and beautiful Beech Fern Phegopteris connectilis. This is a rather uncommon species in the North East but one which I have been encountering more frequently thanks to my ongoing Big Fern Challenge. Now seemed like a good time for a closer look.
Beech Fern
Phegopteris connectilis
Beech Fern is not a species many of us will encounter regularly – unless you’re lucky enough to live in parts of Scotland or perhaps Wales.
This delicate and attractive fern tends to grow in special habitats, including ancient woodlands dominated by Sessile Oak and on shaded, humid gully sides where base-rich water seeps through. Locally, I have found it in both habitats, though the greatest concentrations are nestled among boulder scree in upland areas or tucked away in permanently damp, humid spots around waterfalls and burns. Beech Fern is known to be intolerant of grazing and is therefore often found in places out of reach of passing sheep or deer.
Surprisingly, it is seldom found under Beech trees as it avoids overly acidic substrates, favouring instead neutral to mildly acidic.
Upon further reading, I discovered that Beech Fern has a rather interesting history. It was first reported by Bobart as far back as 1699, who – using typically lengthy Latin – named it Filix minor Britannica pelliculo pallidore alis inferioribus deorsum spectantius. Quite a mouthful, but the name roughly translates to “a small British fern with pale green fronds and downward-pointing pinnae.” Even today, this description remains remarkably accurate!
The name Phegopteris connectilis is a combination of Greek and Latin terms. Phegopteris comes from the Greek words phegos meaning “Beech” and pteris meaning “fern”, describing a fern associated with beech trees. The specific epithet connectilis is Latin and translates “fastened together”.
Although uncommon in lowland areas, the distribution of Beech Fern is considered stable by the BSBI, with new sites being discovered on the edges of its Scottish and Welsh populations. While it may have declined slightly in England, it can still be found in most of its historic haunts.
In terms of its associates, Beech Fern commonly grows alongside Oak Fern Gymnocarpium dryopteris in ancient woodland but in the uplands, may often be found alongside Brittle Bladder-fern Cystopteris fragilis on damp ledges, and in scree alongside Mountain Male-fern Dryopteris oreades.
Identification
Phegopteris connectilis is an easy plant to identify. It is a small, rhizomatous species that can form dense, spreading colonies in suitable habitats. Telling it apart from other ferns shouldn’t be a problem if you focus on a few key features.
Firstly, unlike any other British fern, the pinnae are winged – meaning they’re connected to the rachis along their entire length. This is the origin of the specific epithet connectilis.
Secondly, take a close look at the lowest pair of pinnae. These are opposite, backward-facing, and angled at about 90° to each other. They typically stand erect – resembling a pair of rabbit ears!
That’s all there is to it with this one.
Distribution
In the North East, Beech Fern is an uncommon species. As the orange dots indicate, it hasn’t been recorded at many of its historic sites since the turn of the millennium.
Nevertheless, the species persists in several locations, with notable clusters in the Kielder and Cheviot areas of Northumberland, as well as in the North Pennines of County Durham. It can also be found along watercourses and in remnants of ancient woodland along the River Tyne, from Haltwhistle to Hexham.
Given the lack of recent records, it seems reasonable to conclude that Beech Fern has been lost from most lowland areas of the region.
