Specialist Woodland Grasses

A quick guide to some specialist woodland grasses

I have been meaning to start a series on the UK’s grass species for some time. After a few grass-focused outings this spring, I have decided now is the time. This post, and those to follow, aim to make grass identification a little less daunting for beginners by offering identification tips, quick field clues and habitat notes to help you identify grasses with confidence.

Over time, I will add more pages covering different habitats and grass groups as I continue learning myself, but for now, let’s take a look at some specialist woodland grasses. As ever, all images included are my own unless otherwise stated.

Wood Melick Melica uniflora

Wood Melick Melica uniflora is a dainty and rather beautiful grass of well-managed, typically ancient woodland. Flowering from spring into early summer, it often forms dense tufted patches through rhizomatous spread.

It is a distinctive species, recognised by its graceful sprays of bead-like flowers, ranging from brown to purplish, borne at the tips of fine, branching stems. Confusion is most likely with Mountain Melick Melica nutans, though that species is very rare in much of the UK. To be certain, look for the distinctive bristle pressed against the stem opposite the leaf ligule. The flowers of Wood Melick are also borne on several branches, unlike the next species.

Bristle on the opposing side to the ligule
Delicate, bead-like spikelets
Dainty, branched flowerheads

Mountain Melick Melica nutans

This is a rather rare and presumably declining grass of woodland overlying limestone. You are rather unlikely to come across it in your average woodland, but if you do, good for you!

Altogether rarer than Wood Melick and rather more striking in appearance, Mountain Melick Melica nutans is typically found in woodland over limestone. It bears a one-sided spike of relatively large flowers which, unlike those of M. uniflora, hang delicately from the main stem rather than from side branches. This species also lacks the distinctive bristle or barb at the base of the culm leaves.

Note the absence of the bristle seen in M. uniflora
Larger spikelets than M. uniflora
One-sided spike

Wood Millet Milium effusum

Another grass with delicate, airy, bead-like spikelets, though quite different to the previous two. Wood Millet Milium effusum is a tall woodland grass of damper soils, recognised by its broad, dull green, hairless leaves and rather prominent ligule.

Its tiny, one-flowered spikelets are borne towards the tips of long, drooping branches, creating an elegant, open panicle. The glumes are as long as the entire spikelet and are green with a pale margin, as shown in the image below. Altogether, it is one of the more distinctive woodland grasses.

Prominent ragged ligule
One-flowered spikelets on long, wiry branches
Glumes green with a pale margin

Wood Fescue Drymochloa sylvatica

A rather rare and quite lovely grass, this one has its heartlands in Scotland, Wales and the north of England. Its niche habitat means it is unlikely to be encountered in most woodlands.

A grass of moist, shaded cliffs, ravines and overhangs within ancient woodland, often close to rivers, Wood Fescue Drymochloa sylvatica is a humidity-loving species, typically growing in inaccessible places where competition from other plants is limited.

It forms dense tussocks of flat, shiny evergreen leaves which arch upwards at around 45 degrees, drooping only slightly at the tips. Looking into the tussock, the leaf sheaths are a rich dark brown to purplish colour, lending the base of the plant a distinctive appearance.

Unlike other large woodland fescues, it lacks clasping auricles, while the flowers are unawned. The leaves are also entirely hairless, further helping to separate it from similar woodland grasses.

Delicate, unawned spikelets
Lacks clasping auricles
Leaves arch upwards at around 45 degrees

Wood Barley Hordelymus europaeus

A tall, short-lived grass of woodland over calcareous soils, Wood Barley Hordelymus europaeus is one of our scarcest woodland grasses and has undergone a steady decline across much of its range.

Fortunately, it is the only true barley-like grass likely to be encountered in woodland, and its flower spikes are rather distinctive. Looking more closely, the whole plant is coarsely hairy and bears prominent clasping auricles which project more or less horizontally. In the absence of flowers, this helps separate it from similarly hairy species of Bromopsis. The basal sheaths are also brown rather than reddish, another useful distinction from Bromopsis.

Barley-like flowerheads
Coarsely hairy stems
Prominent clasping auricles, pointed horizontally

Giant Fescue Schedonorus giganteus

A rather tall perennial grass of shady spots within ancient and well-managed woodland, Giant Fescue Schedonorus giganteus is a robust tufted species, often reaching 150 cm in height. It has bright green leaves and distinctive clasping auricles with a purplish tinge, a feature unique among our larger woodland grasses.

The leaves are virtually hairless, tapering gradually to a fine point, while the ligule is short and fairly unobtrusive. Its flower spikes have a distinctive untidy appearance, owing to their long, wavy awns.

Clasping auricles with a purplish tinge
Long-awned spikelets © Mark Welfare
Broad, hairless, bright green leaves

Hairy-brome Bromopsis ramosa

One of the commoner woodland grasses, Hairy-brome Bromopsis ramosa is a species of shaded corners and heavier soils. It is a distinctive tall grass with arching flowerheads, densely hairy sheaths, variably hairy leaves and a prominent ragged ligule.

The lemmas bear long awns and, importantly, the drooping flowerheads usually have just two branches at the lowest node. A rarer related species, Lesser Hairy-brome Bromopsis benekenii (not yet encountered by the author) typically has 3–5 branches at the lowest node instead.

Downward-pointing auricles
Densely hairy sheaths
Graceful, drooping flowers

False Brome Brachypodium sylvaticum

A common grass of shaded woodland, wooded banks and other habitats on heavier soils, False Brome Brachypodium sylvaticum is a clump-forming species which develops distinctive pale green, drooping tussocks.

The leaf sheaths are hairy, at least towards their lower portions, while the ligule is relatively long and pointed. Its flower spikes are also distinctive, initially held upright before arching gracefully over, and are composed of long, slender, overlapping spikelets arranged alternately along the stem.

Leaf sheaths hairy in their upper portion
Pale green, drooping tussocks
False Brome Brachypodium sylvaticum © David Jarema

Cupola Bridge, Stawardpeel & Briarwood Banks

The wooded banks of the River Allen are one of the few places I find myself returning to year after year in search of wild plants (we all have those places, don’t we?). Typically, we follow a familiar route from the National Trust car park at Allen Banks as far as Briarwood Banks, before turning back satisfied with whatever the day has offered.

This spring, though, I was keen to try something a little different. Starting instead from the far end of the gorge at Cupola Bridge, we set out to explore some of the quieter wooded corners we’d somehow overlooked on previous visits. Our usual trips here come a little later in the season, often in search of Bird’s-nest Orchid Neottia nidus-avis, but there was no reason to think an earlier visit would be at all dull.

Cupola Banks

Decanting at Cupola Bridge, the first stretch of our walk saw Matt and me enjoying a leisurely hour exploring Cupola Banks. It is surely one of the finest fragments of ancient woodland in the area, with a reputation for harbouring several local specialities, including Mountain Melick Melica nutans and Wood Barley Hordelymus europaeus. Spoiler: we missed both, though in fairness, we were probably a little early for the barley.

Among the highlights here was Wood Fescue Drymochloa sylvatica, a particularly elegant grass of humid cliffs, shaded overhangs and steep banks where competition from the ever-present Great Wood-rush Luzula sylvatica is reduced. We encountered several fine clumps along this opening stretch, pausing to inspect tussocks for the tell-tale purple-tinged stem bases. Scarce and seemingly in decline, this species is something of a specialist of steep-sided woodlands such as this, often confined to the most inaccessible ledges, making any meaningful monitoring a challenge.

Elsewhere, the supporting cast of woodland grasses was equally pleasing, with plenty of Wood Melick Melica uniflora, alongside Giant Fescue Schedonorus giganteus and Hairy-brome – all good indicators of long-established woodland habitats.

On the floral front, the path edges and wooded slopes were rich with interest. Yellow Pimpernel Lysimachia nemorum, Water Avens Geum rivale and Opposite-leaved Golden-saxifrage Chrysosplenium oppositifolium were all present in good quantity, joined by welcome splashes of colour from Wood Forget-me-not Myosotis sylvatica, Bitter-vetch Lathyrus linifolius and a supporting cast of familiar woodland species.

Less eye-catching, perhaps, but no less noteworthy, were the lingering remains of last season’s Bird’s-nest Orchid. A reminder that we’d arrived a little early, and that a return visit later in the month will be well worth pencilling in.

Of course, I can scarcely go anywhere these days without giving due attention to the ferns (a blessing or a curse, depending on your perspective), and the steep, wooded banks at Cupola proved well worth investigating. The usual suspects were quickly accounted for including Male-fern Dryopteris filix-mas and Lady-fern Athyrium filix-femina, while a few fine clumps of Borrer’s Male-fern Dryopteris borreri were a welcome bonus. Hard Shield-fern Polystichum aculeatum and Polypody Polypodium vulgare were also present, though both were to be expected.

The real highlight came a little higher up the slope, where we were delighted to find neighbouring populations of Beech Fern Phegopteris connectilis and Oak Fern Gymnocarpium dryopteris. Both are scarce locally, and it is always a pleasure to encounter them, particularly growing side by side as they so often do in the damp, humid, rocky woodlands they favour.

To round things off, a small, shaded hollow beside the path held a modest colony of Wood Horsetail Equisetum sylvaticum, its delicately branched foliage easily separating it from other species liable to occur.

Stawardpeel Wood

The next leg of our walk carried us through Stawardpeel Wood, tracing the banks of the Allen and skirting the edges of a sizeable clearfell area. Botanically, much was familiar, with many of the same woodland species continuing along the route. One plant, however, did give us pause: a particularly handsome Dryopteris which, on closer inspection, revealed itself as Western Scaly Male-fern Dryopteris affinis subsp. affinis. A satisfying find, and well worth a closer look.

The more familiar species were in particularly good form along this stretch. Dog-violet Viola riviniana was abundant, joined by swathes of Ramsons Allium ursinum and patches of Wild Strawberry Fragaria vesca – all classic woodland plants.

A final point of interest before reaching Plankey Mill came in the form of a robust member of the pea family scrambling through scrub at the edge of the clearfell. This proved to be Wood Vetch Vicia sylvatica, a Rare Plant Register species in South Northumberland, though on this occasion it was still some way from flowering.

Plankey Mill & Briarwood Banks

The final leg of our small expedition proved to be the most rewarding. Plankey Mill itself offered little of note, aside from a patch of what was likely Upland Enchanter’s-nightshade Circaea x intermedia (one to revisit later in the season), but nearby Briarwood Banks and the pockets of calaminarian grassland in the vicinity were far more engaging.

Venturing back into the woodland, we made a point of checking on a small colony of Early-purple Orchid Orchis mascula that had delighted us on previous visits. Encouragingly, two plants had escaped the attentions of passing feet and were still standing. Nearby, Bugle Ajuga reptans formed carpets in the shade, while a thriving population of Wood Stitchwort Stellaria nemorum was just coming into flower – another northern speciality.

Close to Briarwood Banks, several pockets of calaminarian grassland have developed on soils contaminated by heavy metals and other less savoury residues. These unusual habitats are a joy to visit early in the season thanks to the abundance of Mountain Pansy Viola lutea, which flowers here in April and May.

A short detour brought us to one such clearing, where the pansies were at their peak, forming a kaleidoscope of colour across the turf. Try as we might, we couldn’t pick out any of the pure yellow forms (a personal favourite), but with almost every other variation on display, it was hard to feel disappointed.

And that was that, a thoroughly rewarding spring wander in what must surely be one of the most scenic and botanically rich woodlands in Northumberland. There’s clearly plenty more to come here as the season unfolds, and I’ll have to make a point of returning to Cupola later in the year to seek out some of the species that were still waiting in the wings.